1983 B.A., summa cum laude, Biology and Mathematics, Colby College, Waterville, ME
1983-84 Graduate studies in Applied Mathematics, Rutgers University, New Brunswick, NJ
1985 M.S., Biomathematics, University of California, Los Angeles
1988 Ph.D., Biomathematics (Kenneth Lange, mentor), University of California, Los Angeles
1989-90 Postdoctoral training (Jurg Ott, mentor), Columbia University, New York, NY
Fellowships:
1982 Research Training Program, The Jackson Laboratory, Bar Harbor, ME (David E. Harrison, mentor)
1983 Summer Student Fellow, Woods Hole Oceanographic Institution, MA (Hal Caswell, mentor)
1986-87 Genetic Mechanisms Training Program, UCLA
1987-88 Systems and Integrative Biology Training Program, UCLA
1989-90 Postdoctoral Fellow with Jurg Ott, Ph.D., Columbia University